It's important to distinguish between the original Toho Godzilla films and the more recent Monsterverse films from Legendary Pictures. They have different origin stories. Here's a breakdown: 

Original Toho Films: The 1954 film "Godzilla" (Gojira) established the origin of Godzilla as a creature mutated by nuclear testing. So the original origin is that he is a byproduct of nuclear radiation. 

Monsterverse Films: In the Monsterverse, the origin is changed. Films like "Godzilla" (2014) and "Godzilla: King of the Monsters" (2019), along with supplemental materials, establish that Godzilla is an ancient Titan. These films depict Godzilla as a prehistoric creature that has existed for millions of years, acting as a natural force that maintains balance in the Earth's ecosystem. Also the series "Monarch: Legacy of Monsters" further expands on the knowledge of Godzilla's ancient origins. 

Therefore, while the original 1954 film shows a nuclear-induced origin, the Monsterverse films portray Godzilla as an ancient, natural entity.
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